The Graveyard Gang's peaceful home in a cemetery is threatened when the Tornado Troop scavengers trespass, leading to a scrap between alpha males. Meanwhile, females come into season for a single day, and a vulnerable youngster encounters a predator.
An insight into the Tornado Troop family of ring-tails, and the constant battle for dominance both internally and with the other gang. Unlike the Graveyards, they are a dysfunctional and self-centred bunch, and a five-month-old baby lemur is left to lead a life blighted by loneliness, violence and neglect that could end in tragedy.
The Graveyard Gang faces three rival male lemurs looking for mates, and Blake, the troop's dominant male, prepares for battle. In the chaos that follows, the ring-tails fail to notice six-month-old Hogarth wandering off alone, becoming lost and facing the prospect of a night alone in the wilderness.
The Tornado family faces a crisis when their territory is invaded by farmers. They are forced to go in search of new land, but in the chaos of the move, cub Chelsea is separated from her mother and finds herself straying dangerously close to the tractors. Meanwhile, the mating season gets under way as the males face tough competition to woo the females.
The lemurs face the hardship of the Madagascan winter, and tensions grow among the Tornado Troop. Group leader Electra is forced into exile by aggressive subordinate Amazon, and with a baby on the way and her position increasingly isolated, her survival depends on the mercy of her former gang.
A drought tests the endurance of the lemurs, stretching relations within the families to breaking point. The Graveyard Gang struggles to find nourishment, prompting leader and new mum Crystal to lead a risky foraging expedition, but while they are away the Tornado Troop decides to raid their territory, only to face a tense confrontation when the others return early.
The dry season takes its toll on the Graveyard Gang, putting the lives of Liberty and her newborn in danger as they suffer the effects of malnutrition. Desperation forces the group to undertake the perilous task of foraging through dustbins at a local tourist camp, but the other troops prove an even greater threat to the family's young.
The two families struggle for survival as the winter drought intensifies, and with resources scarce they find themselves clashing over valuable territory. With the groups at war, exiled Tornado Troop leader Electra makes a desperate attempt to join the rival Graveyard Gang, putting herself at risk when the confrontation heats up.
Madagascar continues to face its worst drought in decades, but as two troops resume their desperate search for food, the Graveyard Gang find themselves wandering far from home. Stranded overnight in unfamiliar territory and forced to flee a forest fire, the family face a crisis as one member goes missing.
The Graveyard Gang's territory is invaded by people, which leads to the pack becoming involved in battle after unwittingly moving on to the Tornado Troop's turf. Dominant females Crystal and Amazon go head to head in a fight for supremacy.