Baron Fang resurrects Smilodon the sorceress and ancient mother of the Tusk Tribe in order to destroy the Kamen Riders. Taking on the name Ayanokoji she delivers a voodoo doll that poisons Hongo Takeshi who is taken to a hospital. When Kazami Shiro rushes to the scene she attacks him as Primordial-Tiger, who manages to subdue V3 and locks him up with Hongo. She then targets Ichimonji Hayato… can Hayato free his comrades and best Smilodon?
